<style>
/* =============================================================== *
 * MITO — Mitochondrial Engine · Cellular Energy Stack
 * =============================================================== */
:root {
  --cytoplasm: #f6f0fa;
  --matrix: #ece2f5;
  --membrane: #d8cce8;
  --inner-membrane: #8c64b4;
  --inner-membrane-deep: #5a3a85;
  --cristae-gold: #e8c068;
  --atp-cyan: #00f0d0;
  --myoglobin: #e87a8d;
  --nucleus-blue: #5a7fc4;
  --nucleus-deep: #3a548a;
  --phospholipid: #fff8e0;
  --proton: #b06aff;
  --electron: #3a8fff;
  --carbon: #3a3a3a;
  --oxygen: #d83a2a;
  --phosphate: #f0c020;
  --text-dark: #2a1f3d;
  --text-body: #4a3a5d;
  --text-muted: #7864a0;
  --ff-head: "Quicksand", "Varela Round", "Segoe UI", system-ui, sans-serif;
  --ff-body: "Inter", "Segoe UI", system-ui, sans-serif;
  --ff-mono: "JetBrains Mono", "Consolas", "Courier New", monospace;
  --bio-border: 2px solid rgba(140, 100, 180, 0.3);
  --bio-shadow: 0 8px 24px rgba(120, 60, 160, 0.18);
  --bio-shadow-lg: 0 12px 36px rgba(120, 60, 160, 0.28);
  --membrane-bg: radial-gradient(circle at 50% 0%, #ece2f5, #f6f0fa);
  --cristae-texture: repeating-linear-gradient(90deg, transparent 0 4px, rgba(140, 100, 180, 0.06) 4px 5px);
}

* { box-sizing: border-box; }
html { scroll-behavior: smooth; }
body {
  margin: 0; background: #f6f0fa; color: var(--text-dark);
  font-family: var(--ff-body); font-size: 15px; line-height: 1.6;
  -webkit-font-smoothing: antialiased;
  /* faint fluid texture + slowly spreading calcium waves */
  background-image:
    radial-gradient(circle at 12% 18%, rgba(58, 143, 255, 0.06) 0%, transparent 18%),
    radial-gradient(circle at 88% 42%, rgba(0, 240, 208, 0.05) 0%, transparent 22%),
    radial-gradient(circle at 32% 78%, rgba(176, 106, 255, 0.05) 0%, transparent 20%),
    radial-gradient(circle at 70% 88%, rgba(232, 192, 104, 0.04) 0%, transparent 16%);
  overflow-x: hidden;
  animation: cytoplasm-drift 22s ease-in-out infinite;
}
@keyframes cytoplasm-drift {
  0%, 100% { background-position: 0% 0%, 100% 0%, 0% 100%, 100% 100%; }
  50% { background-position: 8% 4%, 92% -4%, -4% 96%, 108% 92%; }
}
h1,h2,h3,h4,h5 { font-family: var(--ff-head); font-weight: 700; line-height: 1.18; margin: 0 0 .5em; color: var(--text-dark); }
h1 { font-size: clamp(28px, 4vw, 48px); letter-spacing: -0.01em; }
h2 { font-size: clamp(22px, 2.6vw, 30px); }
h3 { font-size: clamp(17px, 1.8vw, 21px); }
a { color: var(--inner-membrane-deep); text-decoration: none; transition: color .2s; }
a:hover { color: var(--atp-cyan); }
img { max-width: 100%; display: block; }
p { margin: 0 0 1em; color: var(--text-body); }
.mono { font-family: var(--ff-mono); }

/* ---------- ambient canvas ---------- */
#mito-canvas { position: fixed; inset: 0; width: 100%; height: 100%; z-index: 0; pointer-events: none; opacity: .65; }
#main, .site-header, .site-footer { position: relative; z-index: 1; }

/* ---------- custom scrollbar (vesicles travelling along a microtubule) ---------- */
::-webkit-scrollbar { width: 14px; }
::-webkit-scrollbar-track {
  background: rgba(140, 100, 180, 0.08);
  border-left: 2px solid rgba(140, 100, 180, 0.18);
}
::-webkit-scrollbar-thumb {
  background: radial-gradient(circle at 50% 50%, #00f0d0, #8c64b4);
  border-radius: 50%;
  border: 2px solid rgba(255,255,255,0.4);
  box-shadow: 0 0 8px rgba(0, 240, 208, 0.4);
  background-size: 100% 50%;
  background-repeat: no-repeat;
  background-position: center;
}
::-webkit-scrollbar-thumb:hover { background: radial-gradient(circle at 50% 50%, #e8c068, #e87a8d); }

/* ---------- layout ---------- */
.layout-with-sidebar { display: grid; grid-template-columns: 1fr; max-width: 1340px; margin: 0 auto; gap: 28px; padding: 0 18px; }
@media (min-width: 1000px) { .layout-with-sidebar { grid-template-columns: 1fr 320px; } }
.layout-main { min-width: 0; }
.eyebrow { display:inline-block; font-family: var(--ff-head); font-size: 11px; letter-spacing: .22em; text-transform: uppercase; color: var(--inner-membrane-deep); background: rgba(255,255,255,0.6); padding: 6px 14px; border: var(--bio-border); border-radius: 999px; box-shadow: var(--bio-shadow); }
.section-head { margin: 32px 0 18px; }
.section-head h2 { margin-top: 10px; color: var(--text-dark); }
.section-head p { color: var(--text-body); max-width: 65ch; }
.muted { color: var(--text-muted); }

/* ---------- header ---------- */
.site-header { position: sticky; top: 0; z-index: 50; background: rgba(246, 240, 250, 0.92); backdrop-filter: blur(10px); border-bottom: 2px solid rgba(140, 100, 180, 0.3); box-shadow: var(--bio-shadow); }
.header-inner { display: flex; align-items: center; gap: 16px; max-width: 1340px; margin: 0 auto; padding: 12px 18px; flex-wrap: wrap; }
.brand { display: flex; align-items: center; gap: 14px; }
.brand-mark { width: 52px; height: 52px; filter: drop-shadow(0 3px 6px rgba(120,60,160,0.3)); }
.brand-mark-rotor { transform-origin: 40px 40px; }
.brand-name { font-family: var(--ff-head); font-weight: 700; font-size: 24px; letter-spacing: .12em; display:block; color: var(--inner-membrane-deep); }
.brand-tag { font-size: 10px; letter-spacing: .22em; color: var(--atp-cyan); text-transform: uppercase; font-family: var(--ff-head); font-weight: 600; }
.main-nav { display: flex; gap: 4px; margin-left: auto; flex-wrap: wrap; }
.main-nav a {
  font-family: var(--ff-head); font-size: 12px; font-weight: 600; letter-spacing: .16em;
  padding: 9px 16px; color: var(--text-body); border: 2px solid transparent;
  border-radius: 999px; transition: all .2s; text-transform: uppercase;
}
.main-nav a:hover { background: rgba(255,255,255,0.6); color: var(--inner-membrane-deep); border-color: rgba(140,100,180,0.3); }
.main-nav a.active { background: var(--inner-membrane); color: var(--phospholipid); border-color: var(--inner-membrane-deep); }
.nav-toggle { display:none; font-family: var(--ff-head); font-weight:700; letter-spacing:.14em; background: var(--inner-membrane); color: var(--phospholipid); border:2px solid var(--inner-membrane-deep); padding:8px 14px; cursor:pointer; border-radius:999px; text-transform: uppercase; }
@media (max-width: 820px) {
  .nav-toggle { display: block; margin-left: auto; }
  .main-nav { display: none; width: 100%; }
  .main-nav.open { display: flex; }
}

/* ---------- cristae maze (top hero) ---------- */
.cristae-maze {
  position: relative; margin: 24px 0; padding: 22px;
  background: var(--membrane-bg);
  border: var(--bio-border);
  border-radius: 32px 48px 36px 52px / 40px 32px 48px 36px; /* irregular mitochondrion shape */
  box-shadow: var(--bio-shadow-lg);
  overflow: hidden;
}
/* cristae folds on the deck surface */
.cristae-maze::before {
  content: ""; position: absolute; inset: 0; z-index: 0; opacity: .6; pointer-events: none;
  background:
    repeating-linear-gradient(120deg, transparent 0 14px, rgba(140,100,180,0.08) 14px 16px),
    repeating-linear-gradient(60deg, transparent 0 18px, rgba(232,192,104,0.05) 18px 20px),
    radial-gradient(circle at 18% 22%, rgba(0,240,208,0.12), transparent 36%),
    radial-gradient(circle at 82% 80%, rgba(176,106,255,0.10), transparent 36%);
}
/* inter-membrane granules (small dots) */
.cristae-maze::after {
  content: ""; position: absolute; inset: 6px; z-index: 0; pointer-events: none;
  background-image:
    radial-gradient(circle at 8% 12%, #8c64b4 1.5px, transparent 2px),
    radial-gradient(circle at 92% 18%, #e8c068 1.5px, transparent 2px),
    radial-gradient(circle at 18% 88%, #00f0d0 1.5px, transparent 2px),
    radial-gradient(circle at 88% 92%, #5a7fc4 1.5px, transparent 2px),
    radial-gradient(circle at 50% 6%, #e87a8d 1.5px, transparent 2px),
    radial-gradient(circle at 50% 94%, #b06aff 1.5px, transparent 2px);
  opacity: 0.6;
}
.cristae-grid {
  position: relative; z-index: 1;
  display: grid; grid-template-columns: repeat(3, 1fr);
  gap: 12px; max-width: 880px; margin: 0 auto;
}
@media (max-width: 700px) { .cristae-grid { grid-template-columns: repeat(2, 1fr); } }

/* ATP synthase chamber — a circular rotor with F1 head and central stator */
.atp-synthase {
  position: relative; aspect-ratio: 3 / 4; cursor: pointer;
  background: radial-gradient(circle at 50% 40%, rgba(232,226,245,0.95), rgba(216,204,232,0.85));
  border: var(--bio-border);
  border-radius: 28px 36px 30px 40px;
  box-shadow: inset 0 0 30px rgba(120,60,160,0.15), var(--bio-shadow);
  overflow: hidden;
  transition: transform .3s, box-shadow .3s;
  display: flex; flex-direction: column; align-items: center; justify-content: center;
  padding: 8px;
}
.atp-synthase:hover { transform: translateY(-3px); box-shadow: inset 0 0 50px rgba(0,240,208,0.18), var(--bio-shadow-lg); }
.atp-synthase.proton-influx {
  box-shadow: inset 0 0 60px rgba(232,192,104,0.45), 0 0 24px rgba(232,192,104,0.4), var(--bio-shadow-lg);
  border-color: var(--cristae-gold);
}
/* the rotor — a circular container holding F1 head subunits and central image */
.synthase-rotor {
  position: relative; width: 80%; aspect-ratio: 1 / 1; max-width: 160px;
  border-radius: 50%;
  background: radial-gradient(circle at 50% 50%, rgba(255,255,255,0.85), rgba(216,204,232,0.6) 60%, rgba(140,100,180,0.4));
  box-shadow: 0 0 16px var(--chamber-accent, var(--atp-cyan)), inset 0 0 18px rgba(120,60,160,0.18);
  border: 2px solid rgba(140, 100, 180, 0.35);
}
/* F1 head — six subunit spheres rotating around the rotor edge */
.f1-head {
  position: absolute; inset: -6%; z-index: 2;
}
.f1-head .f1-subunit {
  position: absolute; width: 18%; aspect-ratio: 1; top: 50%; left: 50%;
  border-radius: 50%;
  background: radial-gradient(circle at 35% 30%, var(--chamber-accent, var(--atp-cyan)), rgba(58,40,85,0.85));
  border: 1.5px solid rgba(255,255,255,0.6);
  box-shadow: 0 0 8px var(--chamber-accent, var(--atp-cyan));
  transform: translate(-50%, -50%) rotate(calc(var(--i) * 60deg)) translateY(-46%);
}
/* central stator — holds the fitness image as the rotor's "core" */
.central-stator {
  position: absolute; top: 18%; left: 18%; right: 18%; bottom: 18%;
  border-radius: 50%; overflow: hidden;
  background: radial-gradient(circle, rgba(58,40,85,0.85), rgba(120,60,160,0.6));
  border: 2px solid rgba(255,255,255,0.4);
  box-shadow: inset 0 0 18px rgba(0,0,0,0.4);
  z-index: 1;
}
.central-stator img {
  width: 100%; height: 100%; object-fit: cover;
  opacity: .9; filter: contrast(1.1) saturate(1.1);
}
.central-stator .chamber-glow {
  position: absolute; inset: -4px; border-radius: inherit;
  background: radial-gradient(circle at 50% 50%, rgba(255,255,255,0.25), transparent 65%);
  pointer-events: none; z-index: 2;
}
/* protein tag below the chamber — fluorescent cyan label */
.cyl-label {
  position: relative; z-index: 3; margin-top: 10px;
  background: rgba(255,255,255,0.85);
  padding: 4px 12px; font-family: var(--ff-head); font-size: 10px;
  letter-spacing: .14em; text-transform: uppercase; color: var(--inner-membrane-deep);
  border: 1.5px solid rgba(0,240,208,0.4); border-radius: 999px;
  box-shadow: var(--bio-shadow); white-space: nowrap; max-width: 96%;
  overflow: hidden; text-overflow: ellipsis; text-align: center;
}
.cyl-label b { display:block; font-size: 11px; color: var(--text-dark); }
.cyl-label span { color: var(--atp-cyan); font-family: var(--ff-mono); font-size: 9px; font-weight: 700; letter-spacing: .04em; }
/* ATP output readout — pops up on hover */
.atp-output-readout {
  position: absolute; top: 8px; left: 50%; transform: translateX(-50%);
  z-index: 5; background: var(--cristae-gold); color: var(--text-dark);
  padding: 4px 12px; font-family: var(--ff-mono); font-size: 10px; font-weight: 700;
  border-radius: 999px; box-shadow: 0 4px 12px rgba(232,192,104,0.6);
  opacity: 0; pointer-events: none; white-space: nowrap;
}
.atp-output-readout::before { content: "⚡ "; }
/* ETC complex icons — connectors between chambers */
.etc-complex {
  position: absolute; pointer-events: none; z-index: 2;
  width: 22px; height: 22px;
  border-radius: 50%; background: radial-gradient(circle, var(--inner-membrane), var(--inner-membrane-deep));
  border: 1.5px solid rgba(255,255,255,0.5);
  box-shadow: 0 0 8px var(--proton);
}
.etc-complex::after {
  content: attr(data-label); position: absolute; top: -16px; left: 50%; transform: translateX(-50%);
  font-family: var(--ff-mono); font-size: 8px; color: var(--electron); letter-spacing: .08em;
  text-transform: uppercase; white-space: nowrap;
}
.electron-particle {
  position: absolute; top: 50%; left: 0; width: 6px; height: 6px;
  border-radius: 50%; background: var(--electron);
  box-shadow: 0 0 8px var(--electron); transform: translateY(-50%);
  pointer-events: none;
}
.proton-pulse-dot {
  position: absolute; top: 50%; left: 50%; width: 8px; height: 8px;
  border-radius: 50%; background: var(--proton);
  box-shadow: 0 0 10px var(--proton); transform: translate(-50%, -50%);
  pointer-events: none;
}
.proton-influx-particle {
  position: absolute; top: 50%; left: 50%; width: 5px; height: 5px;
  border-radius: 50%; background: var(--proton);
  box-shadow: 0 0 6px var(--proton); pointer-events: none;
  transform: translate(-50%, -50%);
}

/* ---------- chamber strip ---------- */
.chamber-strip { display:flex; gap:8px; flex-wrap:wrap; margin: 22px 0; }
.chamber-strip a {
  font-family: var(--ff-head); font-size: 11px; font-weight: 600; letter-spacing: .14em;
  padding: 8px 14px; background: rgba(255,255,255,0.7); color: var(--text-body);
  border: var(--bio-border); border-radius: 999px; transition: all .2s; text-transform: uppercase;
}
.chamber-strip a:hover, .chamber-strip a.active { background: var(--inner-membrane); color: var(--phospholipid); border-color: var(--inner-membrane-deep); }

/* ---------- bands ---------- */
.metabolic-band { margin: 36px 0; padding: 22px; background: rgba(255,255,255,0.55); border: var(--bio-border); border-radius: 24px; box-shadow: var(--bio-shadow); position: relative; overflow: hidden; }
.metabolic-band::before {
  content: ""; position: absolute; inset: 4px; pointer-events: none; opacity: 0.18; border-radius: inherit;
  background-image: repeating-linear-gradient(120deg, transparent 0 8px, rgba(140,100,180,0.4) 8px 9px);
}
.stamped-tags-row { display:flex; flex-wrap:wrap; gap:8px; margin: 14px 0; position: relative; z-index: 1; }
.stamped-tag {
  display:inline-flex; align-items:center; gap:6px;
  background: rgba(0,240,208,0.18); color: var(--inner-membrane-deep);
  padding: 7px 14px; font-family: var(--ff-mono); font-size: 11px; font-weight: 700;
  letter-spacing: .08em; text-transform: uppercase;
  border: 1.5px solid rgba(0,240,208,0.5); border-radius: 999px;
  transition: transform .15s, box-shadow .15s;
}
.stamped-tag:hover { transform: translateY(-2px); box-shadow: 0 4px 12px rgba(0,240,208,0.35); }

/* ---------- metabolic mesh (card grid) ---------- */
.metabolic-mesh {
  position: relative; display: grid;
  grid-template-columns: repeat(auto-fill, minmax(280px, 1fr));
  gap: 22px; z-index: 1;
}
/* faint pathway lines in the background */
.metabolic-mesh::before {
  content: ""; position: absolute; inset: 0; pointer-events: none; z-index: 0; opacity: 0.18;
  background-image: url("data:image/svg+xml;utf8,<svg xmlns='http://www.w3.org/2000/svg' viewBox='0 0 200 200'><g stroke='%238c64b4' stroke-width='0.4' fill='none'><path d='M10 30 Q60 10 100 40 T190 30'/><path d='M20 80 Q80 60 120 90 T180 80'/><path d='M30 140 Q90 120 130 150 T190 140'/><circle cx='40' cy='30' r='3' fill='%2300f0d0'/><circle cx='100' cy='40' r='3' fill='%2300f0d0'/><circle cx='120' cy='90' r='3' fill='%23e8c068'/><circle cx='130' cy='150' r='3' fill='%23e87a8d'/></g></svg>");
  background-size: 360px 360px;
}
@keyframes rotor-spin { from { transform: rotate(0); } to { transform: rotate(360deg); } }
@keyframes rotor-spin-reverse { from { transform: rotate(0); } to { transform: rotate(-360deg); } }

/* glucose molecule card — a hexagonal ring with atom spheres at vertices */
.glucose-card {
  position: relative; display: flex; flex-direction: column; align-items: center; gap: 10px;
  padding: 24px 18px 16px; min-height: 320px; cursor: pointer;
  background: radial-gradient(circle at 50% 38%, rgba(255,255,255,0.85), rgba(216,204,232,0.6));
  border: var(--bio-border); border-radius: 24px;
  box-shadow: var(--bio-shadow);
  transition: transform .25s, box-shadow .25s; z-index: 1;
}
.glucose-card:hover { transform: translateY(-4px); box-shadow: var(--bio-shadow-lg); border-color: var(--atp-cyan); }
.glucose-ring {
  position: relative; width: 150px; height: 150px;
}
.glucose-ring::before {
  content: ""; position: absolute; inset: 0;
  clip-path: polygon(50% 0%, 100% 25%, 100% 75%, 50% 100%, 0% 75%, 0% 25%);
  background: radial-gradient(circle at 50% 50%, rgba(232,192,104,0.45), rgba(140,100,180,0.25));
  border: 2px solid rgba(140,100,180,0.4);
  box-shadow: 0 0 16px rgba(232,192,104,0.4);
}
/* 6 atom spheres at hexagon vertices */
.glucose-ring .atom {
  position: absolute; width: 18px; height: 18px; border-radius: 50%;
  top: 50%; left: 50%;
  border: 1.5px solid rgba(255,255,255,0.7);
  box-shadow: 0 0 6px currentColor;
  z-index: 2;
  transform: translate(-50%, -50%) rotate(calc(var(--i) * 60deg)) translateY(-65px);
}
.glucose-ring .atom.carbon { background: #3a3a3a; color: #3a3a3a; }
.glucose-ring .atom.oxygen { background: #d83a2a; color: #d83a2a; }
.glucose-ring .atom.hydrogen { background: #f8f8f8; color: #fff8e0; }
/* central fitness image inside the ring */
.glucose-ring .ring-image {
  position: absolute; top: 25%; left: 25%; right: 25%; bottom: 25%;
  border-radius: 50%; object-fit: cover;
  border: 2px solid rgba(255,255,255,0.6);
  box-shadow: inset 0 0 16px rgba(0,0,0,0.3);
  z-index: 1;
  filter: contrast(1.1) saturate(1.05);
}
.glucose-ring .phosphate-group {
  position: absolute; top: -8px; right: -8px; width: 22px; height: 22px;
  border-radius: 50%; background: radial-gradient(circle at 35% 30%, #f0c020, #b58a3a);
  border: 1.5px solid var(--phospholipid);
  box-shadow: 0 0 12px #f0c020;
  z-index: 3; opacity: 0;
}
.glucose-ring .phosphate-group::after {
  content: "P"; position: absolute; top: 50%; left: 50%; transform: translate(-50%, -50%);
  font-family: var(--ff-mono); font-size: 10px; font-weight: bold; color: var(--text-dark);
}
.glucose-card .energy-wave {
  position: absolute; top: 50%; left: 50%; transform: translate(-50%, -50%);
  width: 100%; height: 100%; border-radius: 50%;
  border: 2px solid var(--atp-cyan);
  pointer-events: none; z-index: 0;
}
.glucose-card .gc-title { font-family: var(--ff-head); font-weight: 700; font-size: 16px; color: var(--text-dark); text-align: center; }
.glucose-card .gc-meta { font-family: var(--ff-mono); font-size: 11px; color: var(--atp-cyan); text-align: center; letter-spacing: .04em; font-weight: 700; }
.glucose-card .gc-excerpt { font-size: 12px; color: var(--text-body); text-align: center; opacity: .85; }
.glucose-card.phosphorylating { border-color: var(--phosphate); box-shadow: 0 0 32px rgba(240,192,32,0.4), var(--bio-shadow-lg); }

/* metabolic pathway map — a chain of arrows and nodes */
.pathway-map {
  position: relative; background: linear-gradient(180deg, rgba(255,255,255,0.85), rgba(216,204,232,0.55));
  border: var(--bio-border); border-radius: 24px;
  padding: 18px; overflow: hidden; min-height: 240px;
  box-shadow: var(--bio-shadow);
}
.pathway-map .pm-title { font-family: var(--ff-head); font-weight: 700; font-size: 17px; color: var(--text-dark); margin-bottom: 4px; }
.pathway-map .pm-meta { font-family: var(--ff-mono); font-size: 11px; color: var(--atp-cyan); letter-spacing: .04em; margin-bottom: 12px; font-weight: 700; }
.pathway-track {
  position: relative; overflow: hidden; height: 92px;
  border-top: 1.5px dashed rgba(140,100,180,0.4); border-bottom: 1.5px dashed rgba(140,100,180,0.4);
  background: rgba(246,240,250,0.4);
}
.pathway-arrows {
  display: flex; gap: 4px; padding: 8px 0; width: max-content;
}
.pathway-arrow {
  position: relative; display: inline-flex; align-items: center;
  width: 130px; height: 76px;
}
.pathway-arrow .arrow-line {
  position: absolute; top: 50%; left: 0; right: 0; height: 2px;
  background: linear-gradient(90deg, var(--atp-cyan), var(--cristae-gold));
  transform: translateY(-50%);
  box-shadow: 0 0 4px var(--atp-cyan);
}
.pathway-arrow .arrow-head {
  position: absolute; right: 0; top: 50%; transform: translateY(-50%);
  width: 0; height: 0;
  border-left: 10px solid var(--cristae-gold);
  border-top: 6px solid transparent;
  border-bottom: 6px solid transparent;
}
.pathway-arrow .enzyme-name {
  position: absolute; top: 0; left: 50%; transform: translateX(-50%);
  font-family: var(--ff-mono); font-size: 9px; color: var(--inner-membrane-deep);
  letter-spacing: .08em; text-transform: uppercase; white-space: nowrap;
}
.pathway-arrow .flux-particle {
  position: absolute; top: 50%; left: 0; width: 8px; height: 8px;
  border-radius: 50%; background: var(--atp-cyan);
  box-shadow: 0 0 8px var(--atp-cyan); transform: translateY(-50%);
}
.pathway-map .pm-node {
  position: relative; width: 110px; height: 76px; flex-shrink: 0;
  border-radius: 50% / 30%;
  background: linear-gradient(180deg, var(--cristae-gold), var(--myoglobin));
  border: 2px solid rgba(255,255,255,0.5);
  overflow: hidden;
  box-shadow: 0 4px 12px rgba(232,192,104,0.4);
}
.pathway-map .pm-node img { width: 100%; height: 100%; object-fit: cover; opacity: .9; }
.pathway-map .pm-node::after {
  content: ""; position: absolute; left: 50%; top: 50%; width: 14px; height: 14px;
  margin: -7px 0 0 -7px; background: var(--phospholipid); border: 1.5px solid var(--inner-membrane);
  border-radius: 50%; box-shadow: 0 0 8px var(--atp-cyan);
}

/* calcium spark quick-news card */
.ca-spark {
  position: relative; display: flex; align-items: center; gap: 14px;
  background: radial-gradient(circle at 16% 50%, rgba(58,143,255,0.22), transparent 40%),
              linear-gradient(90deg, rgba(255,255,255,0.85), rgba(216,204,232,0.55));
  border: var(--bio-border); padding: 14px 18px 14px 60px;
  border-radius: 16px 24px 16px 24px;
  box-shadow: var(--bio-shadow); transition: transform .2s;
  min-height: 70px; overflow: hidden;
}
.ca-spark:hover { transform: translateX(4px); border-color: var(--electron); }
.ca-spark-ring {
  position: absolute; left: 18px; top: 50%; transform: translate(0, -50%);
  width: 36px; height: 36px; border-radius: 50%;
  border: 2px solid var(--electron);
  box-shadow: 0 0 12px var(--electron), inset 0 0 8px rgba(58,143,255,0.5);
  pointer-events: none;
}
.ca-ion {
  position: absolute; left: 24px; top: 50%; transform: translateY(-50%);
  width: 24px; height: 24px; border-radius: 50%;
  background: radial-gradient(circle at 35% 30%, var(--electron), var(--nucleus-deep));
  border: 1.5px solid var(--phospholipid);
  box-shadow: 0 0 12px var(--electron);
  display: flex; align-items: center; justify-content: center;
  font-family: var(--ff-mono); font-size: 7px; font-weight: bold; color: var(--phospholipid);
  z-index: 2;
}
.ca-spark .fs-title { font-family: var(--ff-head); font-weight: 700; font-size: 14px; color: var(--nucleus-deep); }
.ca-spark .fs-meta { font-family: var(--ff-mono); font-size: 11px; color: var(--atp-cyan); letter-spacing: .04em; font-weight: 700; }
.ca-transient-burst {
  position: absolute; top: 50%; left: 50%; transform: translate(-50%, -50%);
  width: 80px; height: 80px; border-radius: 50%;
  border: 2px solid var(--electron);
  pointer-events: none; z-index: 5;
}

/* wide feature card */
.wide-feature { display:grid; grid-template-columns: 1fr; border:var(--bio-border); background: linear-gradient(180deg, rgba(255,255,255,0.9), rgba(216,204,232,0.6)); box-shadow: var(--bio-shadow); overflow:hidden; transition: transform .25s; border-radius: 24px; }
.wide-feature:hover { transform: translateY(-3px); border-color: var(--atp-cyan); box-shadow: var(--bio-shadow-lg); }
@media (min-width: 640px) { .wide-feature { grid-template-columns: 240px 1fr; } }
.wf-img { width:100%; height:100%; min-height: 180px; object-fit: cover; filter: contrast(1.1) saturate(1.05); }
.wf-body { padding: 18px; }
.wf-body h3 { color: var(--text-dark); }
.wf-excerpt { font-size: 14px; color: var(--text-body); }
.wf-meta { font-family: var(--ff-mono); font-size: 12px; color: var(--atp-cyan); letter-spacing: .04em; font-weight: 700; }

/* ---------- sidebar (cellular dashboard) ---------- */
.layout-sidebar { display: flex; flex-direction: column; gap: 20px; }
.side-block { background: rgba(255,255,255,0.7); border: var(--bio-border); box-shadow: var(--bio-shadow); padding: 16px; border-radius: 20px; backdrop-filter: blur(6px); }
.side-block h3, .side-block h4 { font-family: var(--ff-head); font-size: 12px; letter-spacing: .18em; text-transform: uppercase; color: var(--inner-membrane-deep); border-bottom: 2px solid rgba(140,100,180,0.25); padding-bottom: 8px; margin-bottom: 12px; }

/* ATP energy gauge — circular gauge with golden liquid */
.atp-gauge {
  position: relative; width: 100px; height: 110px; display: flex; flex-direction: column; align-items: center; gap: 12px; cursor: pointer;
}
.atp-gauge .gauge-face {
  position: relative; width: 80px; height: 80px; flex-shrink: 0;
  border-radius: 50% 50% 50% 50% / 60% 60% 40% 40%;
  background: radial-gradient(circle, rgba(58,40,85,0.3), rgba(120,60,160,0.2));
  border: 2px solid rgba(140,100,180,0.4);
  box-shadow: inset 0 0 12px rgba(120,60,160,0.2), var(--bio-shadow);
  overflow: hidden;
}
.atp-gauge .gauge-liquid {
  position: absolute; left: 0; right: 0; bottom: 0; height: 0%;
  background: linear-gradient(180deg, rgba(232,192,104,0.95), rgba(240,192,32,0.85));
  box-shadow: inset 0 4px 8px rgba(255,255,255,0.4), 0 0 16px rgba(232,192,104,0.6);
  transition: height .3s;
}
.atp-gauge .gauge-liquid::before {
  content: ""; position: absolute; top: -3px; left: 0; right: 0; height: 6px;
  background: radial-gradient(circle at 30% 50%, rgba(255,255,255,0.7), transparent 60%);
  border-radius: 50%;
}
.atp-gauge .gauge-label { flex: 1; min-width: 0; }
.atp-gauge .gauge-label b { display: block; font-family: var(--ff-head); font-size: 13px; color: var(--text-dark); overflow: hidden; text-overflow: ellipsis; white-space: nowrap; }
.atp-gauge .gauge-label span { font-family: var(--ff-mono); font-size: 11px; color: var(--cristae-gold); font-weight: 700; }
.gauge-row { display: flex; flex-direction: column; gap: 14px; }
.gauge-row a.atp-gauge { display: flex; flex-direction: row; align-items: center; gap: 12px; text-decoration: none; }

/* signal cascade (kinase comments) */
.cascade-tree { list-style: none; padding: 0; margin: 0; display: flex; flex-direction: column; gap: 10px; }
.cascade-kinase {
  position: relative; background: rgba(255,255,255,0.7); border: 2px solid var(--inner-membrane);
  padding: 10px 12px 10px 36px; font-family: var(--ff-mono); font-size: 12px;
  border-radius: 12px; cursor: pointer; transition: all .2s;
  margin-left: calc(var(--depth, 0) * 16px);
}
.cascade-kinase::before {
  content: ""; position: absolute; left: 10px; top: 50%; transform: translateY(-50%);
  width: 16px; height: 16px; border-radius: 50%;
  background: radial-gradient(circle, var(--inner-membrane), var(--inner-membrane-deep));
  border: 1.5px solid var(--phospholipid);
  box-shadow: 0 0 6px var(--inner-membrane);
}
.cascade-kinase.active { border-color: var(--atp-cyan); box-shadow: 0 0 12px rgba(0,240,208,0.4); }
.cascade-kinase.active::before { background: radial-gradient(circle, var(--atp-cyan), var(--nucleus-deep)); }
.cascade-kinase.highlighted { border-color: var(--cristae-gold); box-shadow: 0 0 12px rgba(232,192,104,0.4); }
.cascade-meta { color: var(--inner-membrane-deep); font-size: 10px; letter-spacing: .04em; display: block; font-weight: 700; }
.cascade-text { color: var(--text-body); display: block; margin-top: 2px; }

/* membrane potential voting */
.membrane-vote { position: relative; padding: 8px; }
.membrane-cell {
  position: relative; width: 100%; height: 80px; margin: 14px 0;
  background: radial-gradient(circle, rgba(90,127,196,0.4), rgba(58,84,138,0.6));
  border: 2px solid var(--nucleus-blue);
  border-radius: 50%; overflow: hidden;
  box-shadow: inset 0 0 16px rgba(58,84,138,0.5), var(--bio-shadow);
  transition: all .3s;
  display: flex; align-items: center; justify-content: center;
}
.membrane-cell.depolarized { background: radial-gradient(circle, rgba(216,58,42,0.6), rgba(120,40,30,0.7)); border-color: var(--oxygen); box-shadow: inset 0 0 20px rgba(216,58,42,0.6), 0 0 16px rgba(216,58,42,0.4); }
.membrane-cell.hyperpolarized { background: radial-gradient(circle, rgba(58,143,255,0.6), rgba(20,60,120,0.7)); border-color: var(--electron); box-shadow: inset 0 0 20px rgba(58,143,255,0.6), 0 0 16px rgba(58,143,255,0.4); }
.potential-readout { font-family: var(--ff-mono); font-size: 16px; font-weight: bold; color: var(--phospholipid); text-shadow: 0 0 8px rgba(0,0,0,0.5); }
.vote-row { display: flex; gap: 22px; justify-content: center; margin: 14px 0; }
.vote-side {
  position: relative; padding: 10px 16px; cursor: pointer;
  background: rgba(255,255,255,0.7); border: 2px solid rgba(140,100,180,0.3);
  border-radius: 12px; font-family: var(--ff-head); font-size: 12px; font-weight: 700;
  letter-spacing: .12em; text-transform: uppercase; color: var(--text-dark);
  transition: all .15s;
}
.vote-side:hover { transform: scale(1.05); }
.vote-side[data-dir="depolarize"]:hover { border-color: var(--oxygen); color: var(--oxygen); }
.vote-side[data-dir="hyperpolarize"]:hover { border-color: var(--electron); color: var(--electron); }
.ion-splash { position: absolute; width: 6px; height: 6px; border-radius: 50%; pointer-events: none; left: 50%; top: 50%; }

/* calcium oscillation calendar */
.calcium-calendar {
  position: relative; height: 120px; margin: 8px 0;
  background: rgba(58,40,85,0.15); border: 2px solid rgba(140,100,180,0.3);
  box-shadow: inset 0 0 14px rgba(120,60,160,0.2);
  border-radius: 16px; overflow: hidden;
}
.calcium-curve-svg { position: absolute; inset: 0; width: 100%; height: 100%; }
.calcium-cursor {
  position: absolute; width: 14px; height: 14px; border-radius: 50%;
  background: radial-gradient(circle, var(--electron), var(--nucleus-deep));
  border: 2px solid var(--phospholipid);
  box-shadow: 0 0 12px var(--electron); z-index: 3;
  transform: translate(-50%, -50%);
}
.calcium-today {
  position: absolute; width: 2px; top: 0; bottom: 0;
  background: var(--atp-cyan); z-index: 2;
  box-shadow: 0 0 8px var(--atp-cyan);
}
.ca-wave-pulse {
  position: absolute; top: 0; bottom: 0; width: 30px;
  background: linear-gradient(90deg, transparent, rgba(58,143,255,0.4), transparent);
  z-index: 1; pointer-events: none;
}
.calcium-axis-x { font-family: var(--ff-mono); font-size: 9px; color: var(--text-muted); letter-spacing: .12em; text-transform: uppercase; text-align: center; margin-top: 6px; }
.scent-hint { font-size: 11px; color: var(--text-muted); margin-top: 8px; font-style: italic; text-align: center; }

/* ---------- pagination ---------- */
.pagination { display:flex; gap: 6px; justify-content: center; margin: 28px 0; flex-wrap: wrap; }
.pagination a, .pagination span { font-family: var(--ff-head); font-weight: 700; padding: 8px 14px; border: var(--bio-border); background: rgba(255,255,255,0.7); color: var(--text-body); border-radius: 999px; }
.pagination a:hover { background: var(--inner-membrane); color: var(--phospholipid); }
.pagination .current { background: var(--inner-membrane); color: var(--phospholipid); border-color: var(--inner-membrane-deep); }

/* ---------- detail page ---------- */
.detail-breadcrumb { font-family: var(--ff-mono); font-size: 12px; color: var(--text-muted); margin-bottom: 16px; display:flex; gap:8px; flex-wrap: wrap; align-items: center; letter-spacing: .08em; }
.detail-breadcrumb a { color: var(--inner-membrane-deep); }
.detail-hero { display: grid; grid-template-columns: 1fr; gap: 22px; margin-bottom: 24px; }
@media (min-width: 760px) { .detail-hero { grid-template-columns: 340px 1fr; } }
.detail-hero-frame { position: relative; border: var(--bio-border); box-shadow: var(--bio-shadow-lg); overflow: hidden; background: var(--membrane); border-radius: 24px; }
.detail-hero-frame::before { content: ""; position: absolute; inset: 0; z-index: 2; pointer-events: none; background: repeating-linear-gradient(120deg, transparent 0 8px, rgba(140,100,180,0.12) 8px 9px); border-radius: inherit; }
.detail-hero-img { width: 100%; height: 380px; object-fit: cover; filter: contrast(1.1) saturate(1.05); }
.detail-hero-glow { position:absolute; inset:0; background: radial-gradient(circle at 30% 30%, rgba(0,240,208,0.25), transparent 60%); pointer-events: none; z-index: 3; }
.detail-shape { display:inline-block; font-family: var(--ff-head); font-size: 11px; letter-spacing: .18em; text-transform: uppercase; color: var(--phospholipid); background: var(--inner-membrane); padding: 6px 14px; border-radius: 999px; box-shadow: var(--bio-shadow); }
.detail-stats { display: flex; gap: 10px; flex-wrap: wrap; margin: 14px 0; }
.stat { background: rgba(255,255,255,0.7); border: var(--bio-border); padding: 8px 14px; font-family: var(--ff-mono); font-size: 11px; color: var(--text-muted); border-radius: 12px; }
.stat b { font-family: var(--ff-head); font-size: 16px; color: var(--cristae-gold); }
.detail-tags { display: flex; gap: 6px; flex-wrap: wrap; }
.detail-tag { font-family: var(--ff-mono); font-size: 10px; padding: 4px 10px; background: rgba(0,240,208,0.18); color: var(--inner-membrane-deep); border: 1.5px solid rgba(0,240,208,0.4); border-radius: 999px; letter-spacing: .08em; text-transform: uppercase; font-weight: 700; }
.detail-body { display: grid; grid-template-columns: 1fr; gap: 24px; }
@media (min-width: 1000px) { .detail-body { grid-template-columns: 1fr 320px; } }
.detail-content h2 { margin-top: 18px; color: var(--text-dark); }
.detail-desc { font-size: 16px; line-height: 1.75; color: var(--text-body); }
.atp-note { background: linear-gradient(180deg, rgba(255,255,255,0.85), rgba(216,204,232,0.55)); border: var(--bio-border); box-shadow: var(--bio-shadow); padding: 18px; margin: 18px 0; border-radius: 20px; }
.atp-coord { font-family: var(--ff-mono); font-size: 11px; letter-spacing: .14em; text-transform: uppercase; color: var(--atp-cyan); font-weight: 700; }
.atp-text { font-size: 14px; color: var(--text-body); margin: 8px 0; font-style: italic; }
.atp-sign { font-family: var(--ff-mono); font-size: 11px; color: var(--text-muted); letter-spacing: .08em; }
.detail-related h3 { font-family: var(--ff-head); margin: 24px 0 12px; color: var(--text-dark); }

/* ---------- static pages ---------- */
.static-hero { margin-bottom: 24px; }
.static-lede { font-size: 17px; color: var(--text-body); max-width: 65ch; font-style: italic; }
.static-card { background: rgba(255,255,255,0.75); border: var(--bio-border); box-shadow: var(--bio-shadow); padding: 20px; margin-bottom: 16px; border-radius: 20px; }
.static-card h2 { color: var(--text-dark); }
.layer-list { list-style: none; padding: 0; }
.layer-list li { padding: 8px 0; border-bottom: 1px dashed rgba(140,100,180,0.3); color: var(--text-body); }
.layer-list li b { color: var(--inner-membrane-deep); }
.transcription-form, .spore-form { display: flex; flex-direction: column; gap: 10px; }
.transcription-form label, .spore-form label { display: flex; flex-direction: column; gap: 4px; font-size: 12px; font-family: var(--ff-head); letter-spacing: .14em; text-transform: uppercase; color: var(--text-body); }
.transcription-form input, .spore-form input, .spore-form textarea { padding: 10px; border: var(--bio-border); background: rgba(255,255,255,0.85); color: var(--text-dark); font-family: var(--ff-mono); font-size: 14px; border-radius: 12px; }
.transcription-form input:focus, .spore-form input:focus { outline: 2px solid var(--atp-cyan); }
.transcription-form button, .spore-form button { align-self: flex-start; padding: 12px 22px; background: var(--inner-membrane); color: var(--phospholipid); border: 2px solid var(--inner-membrane-deep); font-family: var(--ff-head); font-weight: 700; letter-spacing: .14em; cursor: pointer; box-shadow: var(--bio-shadow); text-transform: uppercase; border-radius: 999px; }
.transcription-form button:hover, .spore-form button:hover { transform: translateY(-2px); background: var(--atp-cyan); color: var(--text-dark); }
.static-aside { background: rgba(255,255,255,0.7); border: var(--bio-border); padding: 18px; margin-top: 16px; border-radius: 20px; }

/* ---------- search ---------- */
.search-box { display: flex; align-items: center; gap: 8px; background: rgba(255,255,255,0.85); border: var(--bio-border); box-shadow: var(--bio-shadow); padding: 8px 16px; border-radius: 999px; }
.search-box input { flex: 1; border: none; background: transparent; font-family: var(--ff-mono); font-size: 14px; padding: 6px; outline: none; color: var(--inner-membrane-deep); }
.search-count { font-family: var(--ff-mono); font-size: 11px; color: var(--text-muted); }
.search-results { display: grid; grid-template-columns: repeat(auto-fill, minmax(240px,1fr)); gap: 12px; margin-top: 16px; }
.search-cell { display: flex; gap: 10px; background: rgba(255,255,255,0.75); border: var(--bio-border); padding: 10px; box-shadow: var(--bio-shadow); border-radius: 16px; }
.search-cell .sc-thumb { width: 56px; height: 64px; flex-shrink: 0; overflow: hidden; border: 1.5px solid rgba(140,100,180,0.3); border-radius: 12px; }
.search-cell .sc-thumb img { width:100%; height:100%; object-fit: cover; }
.search-cell .sc-meta { display:block; font-family: var(--ff-mono); font-size: 11px; color: var(--atp-cyan); letter-spacing: .04em; font-weight: 700; }
.search-cell .sc-body b { color: var(--text-dark); font-family: var(--ff-head); }

/* ---------- footer (nucleus & gene bank) ---------- */
.site-footer { position: relative; margin-top: 60px; background: radial-gradient(ellipse at center, #d8cce8 0%, #a09abc 100%); border-top: 2px solid rgba(140,100,180,0.4); overflow: hidden; }
/* nuclear pore complex — small octagonal dots on the nuclear envelope */
.site-footer::before {
  content: ""; position: absolute; inset: 0; z-index: 0; pointer-events: none; opacity: 0.6;
  background-image:
    radial-gradient(circle at 6% 12%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 18% 28%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 32% 14%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 48% 32%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 64% 18%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 78% 36%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 92% 22%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 10% 60%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 28% 76%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 50% 88%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 72% 64%, rgba(255,255,255,0.7) 2px, transparent 3px),
    radial-gradient(circle at 88% 80%, rgba(255,255,255,0.7) 2px, transparent 3px);
}
.nucleus-floor { position: relative; padding: 36px 18px 28px; overflow: hidden; }
.nucleus-grid { position: relative; z-index: 1; display: grid; grid-template-columns: repeat(auto-fill, minmax(170px, 1fr)); gap: 18px; max-width: 1180px; margin: 0 auto; }

/* gene locus card — a Giemsa-banded chromosomal band */
.gene-locus {
  position: relative; aspect-ratio: 1 / 1.1; cursor: pointer;
  background: radial-gradient(circle at 50% 60%, rgba(140,100,180,0.25), transparent 65%),
              linear-gradient(180deg, rgba(216,204,232,0.85), rgba(160,154,188,0.7));
  border: var(--bio-border); box-shadow: var(--bio-shadow);
  border-radius: 16px; overflow: hidden; display: flex; align-items: flex-end; justify-content: center;
  padding: 14px; transition: all .25s;
}
.gene-locus:hover { transform: translateY(-3px); border-color: var(--atp-cyan); box-shadow: 0 0 24px rgba(0,240,208,0.4), var(--bio-shadow); }
.gene-locus .locus-image {
  position: relative; z-index: 2; width: 86%; height: 70%; object-fit: cover;
  filter: contrast(1.1) saturate(0.9); opacity: .9;
  border: 2px solid rgba(255,255,255,0.6); border-radius: 12px;
}
/* Giemsa bands — horizontal stripes overlay */
.gene-locus::before {
  content: ""; position: absolute; left: 8%; right: 8%; top: 14%; height: 56%;
  background: repeating-linear-gradient(0deg,
    rgba(58,40,85,0.55) 0 6px,
    rgba(232,192,104,0.55) 6px 10px,
    rgba(58,40,85,0.55) 10px 16px,
    rgba(140,100,180,0.4) 16px 22px,
    rgba(58,40,85,0.55) 22px 28px);
  border-radius: 6px; z-index: 1; opacity: .55;
}
.gene-locus .locus-title {
  position: absolute; bottom: 8px; left: 50%; transform: translateX(-50%);
  font-family: var(--ff-head); font-size: 11px; color: var(--text-dark);
  text-align: center; z-index: 3; padding: 0 6px; letter-spacing: .1em; text-transform: uppercase;
  font-weight: 700;
}

/* telomere clock — TTAGGG repeats shrinking over time */
.telomere-wall { position: relative; z-index: 1; max-width: 980px; margin: 32px auto 0; padding: 18px 20px; background: rgba(58,40,85,0.5); border: 2px solid rgba(140,100,180,0.4); border-radius: 20px; box-shadow: inset 0 0 14px rgba(0,0,0,0.2); }
.telomere-clock { display: flex; align-items: center; gap: 4px; flex-wrap: wrap; }
.telomere-repeat {
  display: inline-block; padding: 4px 8px; font-family: var(--ff-mono); font-size: 10px; font-weight: 700;
  background: rgba(0,240,208,0.25); color: var(--phospholipid); border-radius: 6px;
  border: 1px solid rgba(0,240,208,0.5);
  letter-spacing: .12em;
}
.telomere-crisis {
  display: inline-block; padding: 4px 10px; font-family: var(--ff-head); font-size: 10px; font-weight: 700;
  background: rgba(216,58,42,0.4); color: var(--phospholipid); border-radius: 6px;
  border: 1px solid rgba(216,58,42,0.7); letter-spacing: .14em; text-transform: uppercase;
  margin: 0 6px;
}
.telomere-extend {
  display: inline-block; padding: 4px 10px; font-family: var(--ff-head); font-size: 10px; font-weight: 700;
  background: rgba(232,192,104,0.45); color: var(--text-dark); border-radius: 6px;
  border: 1px solid rgba(232,192,104,0.7); letter-spacing: .14em; text-transform: uppercase;
  margin: 0 6px;
}

.footer-grid { display: grid; grid-template-columns: 1fr; gap: 22px; max-width: 1200px; margin: 0 auto; padding: 28px 18px; position: relative; z-index: 1; }
@media (min-width: 700px) { .footer-grid { grid-template-columns: 1.4fr 1fr 1fr 1.2fr; } }
.footer-col h4 { font-family: var(--ff-head); font-size: 12px; letter-spacing: .18em; text-transform: uppercase; color: var(--inner-membrane-deep); margin-bottom: 10px; border-bottom: 2px solid rgba(140,100,180,0.3); padding-bottom: 6px; }
.footer-col ul { list-style: none; padding: 0; margin: 0; }
.footer-col li { padding: 4px 0; }
.footer-col a { color: var(--text-body); opacity: .9; }
.footer-col a:hover { opacity: 1; color: var(--atp-cyan); }
.mito-logo { display: flex; align-items: center; gap: 12px; margin-bottom: 14px; }
.mito-logo .brand-mark { width: 52px; height: 52px; }
.mito-name { font-family: var(--ff-head); font-weight: 700; font-size: 22px; letter-spacing: .12em; color: var(--inner-membrane-deep); }
.mito-tag { font-size: 10px; letter-spacing: .18em; color: var(--atp-cyan); text-transform: uppercase; font-family: var(--ff-head); font-weight: 600; }

/* transcription subscribe — DNA double helix that unzips */
.transcription-cell {
  position: relative; background: rgba(255,255,255,0.75); border: var(--bio-border); padding: 18px; margin-top: 12px;
  box-shadow: var(--bio-shadow); border-radius: 20px; overflow: hidden;
}
.dna-helix {
  position: relative; height: 50px; margin-bottom: 12px;
  display: flex; align-items: center; gap: 4px;
}
.dna-strand {
  position: absolute; top: 50%; transform: translateY(-50%); height: 4px;
  width: 100%; border-radius: 2px;
}
.dna-strand:nth-child(1) { background: linear-gradient(90deg, var(--atp-cyan), var(--inner-membrane)); transform: translateY(-12px); }
.dna-strand:nth-child(2) { background: linear-gradient(90deg, var(--myoglobin), var(--cristae-gold)); transform: translateY(12px); }
.dna-base {
  position: absolute; top: 50%; transform: translate(-50%, -50%);
  width: 10px; height: 10px; border-radius: 50%;
  background: var(--phospholipid); border: 1px solid var(--inner-membrane);
}
.mrna-strand {
  position: absolute; top: 50%; left: 10%; transform: translateY(-50%);
  height: 3px; width: 30%; border-radius: 2px;
  background: linear-gradient(90deg, var(--atp-cyan), var(--phosphate));
  box-shadow: 0 0 8px var(--atp-cyan);
  opacity: 0; transition: opacity .3s;
}
.mrna-strand.transcribed { opacity: 1; }
.transcription-form input { width: 100%; padding: 10px; border: var(--bio-border); background: rgba(255,255,255,0.9); color: var(--inner-membrane-deep); font-family: var(--ff-mono); margin: 8px 0; font-size: 14px; border-radius: 12px; }
.rna-polymerase-btn {
  display: block; width: 100%; padding: 12px; background: var(--inner-membrane); color: var(--phospholipid);
  border: 2px solid var(--inner-membrane-deep); font-family: var(--ff-head); font-weight: 700; letter-spacing: .14em;
  cursor: pointer; box-shadow: var(--bio-shadow); text-transform: uppercase; border-radius: 999px;
  transition: all .2s;
}
.rna-polymerase-btn:hover { background: var(--atp-cyan); color: var(--text-dark); transform: translateY(-2px); }
.transcription-confirm { background: rgba(0,240,208,0.25); color: var(--text-dark); padding: 14px; margin-top: 8px; font-family: var(--ff-head); font-size: 13px; letter-spacing: .1em; border: 1.5px solid rgba(0,240,208,0.6); text-align: center; border-radius: 12px; }
.transcription-confirm::before { content: "◆ TRANSCRIBED "; color: var(--atp-cyan); font-weight: 700; }

.tagline-line { font-family: var(--ff-head); font-size: 13px; letter-spacing: .16em; text-align: center; padding: 20px; border-top: 2px solid rgba(140,100,180,0.3); border-bottom: 2px solid rgba(140,100,180,0.3); color: var(--text-dark); display: flex; align-items: center; justify-content: center; gap: 12px; text-transform: uppercase; background: rgba(255,255,255,0.5); }
.footer-bottom { text-align: center; padding: 14px; font-family: var(--ff-mono); font-size: 11px; color: var(--text-muted); letter-spacing: .12em; text-transform: uppercase; }

.exit-arrow { position: fixed; right: 22px; bottom: 22px; width: 46px; height: 46px; background: var(--inner-membrane); color: var(--phospholipid); border: 2px solid var(--inner-membrane-deep); display: flex; align-items: center; justify-content: center; cursor: pointer; opacity: 0; pointer-events: none; transition: opacity .3s; box-shadow: var(--bio-shadow-lg); z-index: 60; border-radius: 50%; }
.exit-arrow.show { opacity: 1; pointer-events: auto; }
.exit-arrow svg { width: 22px; height: 22px; }

/* reveal */
[data-reveal] { opacity: 0; transform: translateY(18px); transition: opacity .6s, transform .6s; }
[data-reveal].in { opacity: 1; transform: none; }
@media (prefers-reduced-motion: reduce) { [data-reveal] { opacity:1; transform:none; } }

/* small screens */
@media (max-width: 600px) {
  .metabolic-mesh { grid-template-columns: 1fr; }
  .detail-hero-img { height: 260px; }
  .cascade-kinase { font-size: 11px; }
}
</style>